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They admittedly speculate that the "intensity and duration of the bottleneck(s) leading to breed formation may have been extreme in cKCs relative to in most other breeds." The "bottlenecks" they refer to include (a) development of a brachycephalic variety of the King Charles spaniel, presumably in the late 1800s, followed by "the old, more long-nosed type was restored by a small breeding initiative starting in the 1920's"

2020 Auswirkungen einer L-Carnitin-Supplementierung auf den Gewichtsverlust und die Krperzusammensetzung Yarizadh et al