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10.1172/JCI34827 86

Besides H3K9, multiple lysine and arginine residues of histones (such as H3R2, H3K4, H3K27, H3K79, H4R3, H4K20, and H2BK5) also undergo methylation/demethylation dynamics and exhibit pleiotropic roles in gene transcription (65)

3.2 DNA fragmentation In addition to lipid peroxidation, oxidative stress induces significant damage to sperm DNA, leading to DNA fragmentation ( Figure 2 )