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Brown adipose tissue (BAT), brain (cortex), heart, intestine, kidney, liver, lung, muscle, spleen and white adipose tissue (WAT) were dissected and quickly frozen in liquid nitrogen immediately and stored at 80 C until metabolite extraction

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doi: 10.1016/j.bbi.2019.07.028

Upon ER stress, they are transported to the Golgi where they are proteolytically cleaved and then translocate to the nucleus, where they induce UPR genes